for his work on ribonucleaseandfor their contribution to the understanding of the connection between chemical structure and catalytic activity of the ribonuclease molecule
for their elucidation of the enzymatic mechanism underlying the synthesis of adenosine triphosphateandfor his discovery of an ion-transporting enzyme, Na+, K+ -ATPase
... as intensives[?]/frequentives[?], desideratives[?], causatives[?], and benedictives[?] derived from more basic forms). Each verb is also has a grammatical voice: ...