for his work on ribonucleaseandfor their contribution to the understanding of the connection between chemical structure and catalytic activity of the ribonuclease molecule
for their elucidation of the enzymatic mechanism underlying the synthesis of adenosine triphosphateandfor his discovery of an ion-transporting enzyme, Na+, K+ -ATPase
... now in a form in which we can conveniently complete the square[?]. To "complete the square" is to add a constant (i.e., in this case, a quantity that does not depend on x) ...